Gramma the Galápagos tortoise, oldest resident of San Diego Zoo, dies at about 141 The beloved Galápagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has died Original Source: https://abcnews.go.com/US/wireStory/gramma-galpagos-tortoise-oldest-resident-san-diego-zoo-127847821